The Hawaiian Humane Society on Friday issued a citation on 310 counts of animal cruelty against the owners of a Makaha animal shelter, where more than 300 dogs were seized in October.

The seizures took place at the Friends for Life shelter, where Humane Society representatives said the animals were living in “inhumane” conditions, contaminated with feces and urine. Two owners of the shelter facing counts are David Lanny Moore and June Moore, according to Humane Society spokeswoman Allison Andrade Gammel.

Each count carries a maximum penalty of $2,000 in fines and a year of jail time, Andrade said. The Humane Society is deputized by the Honolulu Police Department to issue those counts, she said.

Representatives for Friends for Life could not immediately be reached Friday.

Dogs seized in October are currently in about 200 foster homes across the island, with others at the Humane Society’s McCully-Moiliili facility, Andrade said. The organization has an ongoing investigation related to those seizures, Andrade said.

In October, some Friends for Life volunteers defended the shelter, saying it is willing to take in scores of abandoned animals while adhering to a no-kill policy.

Andrade said the facility was required by law to give the dogs sufficient care to preserve their well-being at a minimum, but “that was not being done for these animals.”

The Humane Society continues to seek foster families for the dogs seized. Anyone interested is encouraged to call 356-2200 or visit hawaiianhumane.org.